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@Aaron Boranian is perfectly right (and his answer should be accepted).
To state the obvious and be perfectly clear, the rule is:
* anything that starts with !-
is a comment that isn't retained, and,
* anything that starts with !
(no -
) would be retained.
In both cases, these are comments and do not affect in any shape or form how stuff is being understood by the E+ Input processor that reads your IDF file.
These !-
in particular will be (re)added by the IDF editor upon save, and you can think of them as visual aids when you edit stuff in a text editor.